Sauropoda
More commonly known to the public under
the informal name "brontosaurs," sauropods are probably one of the top three
general types of classic dinosaurs most likely to enter the average person's mind when
they hear the word dinosaur (the other two being tyrannosaurids
and "raptors", with ceratopsids
and stegosaurids rounding out the top five). Their
general form is instantly recognizable: four stout legs (interestingly, derived
sauropods reduce the number of bones in their fingers and toes; titanosaurians,
for example, may well have lacked hand claws entirely), long neck and tail, miniscule
head, and large body. Some might consider them boring, because it can be claimed
once you've seen one, you've pretty much seen them all, but they are worth a second
look. They were one of Nature's most amazing feats of living engineering, every
feature a balance between strength and lightness on a scale never seen before or
since. All had at least depressions in
their vertebrae, and in some, like the brachiosaurids,
the verts are heavily excavated (such holes are called pleurocoels) and even the ribs have
hollowing. The largest
sauropod is
almost certainly not known yet. Stereotypically, sauropods lived in bodies of water,
but it has been shown that such a lifestyle would have seriously hindered breathing, to
say the least. Although sometimes restored with an elephant-like hide,
sauropods were scaled and many may have had nonbony spines, like those of an
iguana, running down their backs. In addition, there has been recent
discussion about the orientation of the shoulder blades; if set more level, as
has been suggested, the front end of the skeleton is raised noticeably.
Sauropods appear to come in three or four general varieties: macronarians,
diplodocoids, and a spray of basal forms for variety. Macronarians include the
camarasaurids, brachiosaurids, and titanosaurians, rather
conservative sauropod groups, but ironically the last sauropods appear to have been
titanosaurians. Some forms are known to have possessed body armor. They are foremost a Gondwana
phenomenon. The diplodocoids, on the other hand, are
mainly Laurasian and African in membership, best known for being very long, lean and somewhat lower
to the ground than macronarians.
<--Sauropoda
`--+--+--Antetonitrus
| `--Lessemsaurus
`--+--Chinshakiangosaurus
|--Gongxianosaurus
|--Isanosaurus
|--Kotasaurus
`--Gravisauria
|--Vulcanodon
`--+--+--Spinophorosaurus
| `--Tazoudasaurus
`--+--Barapasaurus
`--+--Rhoetosaurus
`-->Eusauropoda
This looks very short, but trust me here; there are enough dubious sauropods to take up
plenty of space.
Sauropoda:
| Taxon or Taxa: | Time/Place: | Comments: |
| Antetonitrus ingenipes Yates and Kitching, 2003 | early Norian (LTr) of South Africa | A newly described, small but robust basal sauropod, Antetonitrus had been identified as a specimen of Euskelosaurus. It is based on a partial skeleton including some vertebrae and ribs, and most of the fore and hindlimbs. The hands are interesting in that the thumbs appear to retain a manipulatory ability, like the prosauropods. |
| Lessemsaurus sauropoides Bonaparte, 1999 | late Norian (LTr) of Argentina | Based on a partial vertebral column, with fairly tall neural arches, this basal sauropod was originally described as a "melanorosaurid" prosauropod, but seems to be closer to more derived sauropods than Melanorosaurus. |
| Chinshakiangosaurus chunghoensis Ye vide Dong, 1992 | EJ of China | This name had been floating around forever, and was mostly ignored, until suddenly people figured out that it had been published or something. Seriously, this was about as obscure as could be, usually tossed off as an unofficial name for a sauropod or "melanorosaurid". Now, though, the type lower jaw, and some other remains as well (verts, scapulae, pelvic bones, and leg bones), has hit the presses. It is considered some sort of basal sauropod, but things are still exploding violently in that area of dinosaur research, so check back in a couple of years once Prosauropod Pinball is finished. It seems to have had cheeks. |
| Gongxianosaurus shibeiensis He, Wang, Liu S., Zhou, Liu T., Cai, and Dai, 1998 | ?Pliensbachian-Toarcian (EJ) of China | This is another prosauropod-like (?actual prosauropod) EJ sauropod. 3 partial skeletons are known, including cranial material. The tail has simple chevrons which, like those of some prosauropods, are rather long, forming a deep tail. A restoration shows belly ribs, but I have been unable to find them in the description. A second species may be present. |
| Isanosaurus attavipachi Buffetaut, Suteethorn, Cuny, Tong, Le Loeuff, Khansubha, and Jongautchariyakul, 2000 | late Norian or Rhaetian (LTr) of Thailand | One of the earliest-known sauropods, Isanosaurus is based on partial postcranial remains including a cervical, dorsal, six caudals, sternal plate, scapula, and femur, probably from a subadult individual. It appears to have been very basal. A humerus from the same time and place indicate an animal about 12-15 in length, but it is not known if it is the same animal as Isanosaurus. |
| Kotasaurus yamanpalliensis Yadagiri, 1988 | Sinemurian-Pliensbachian (EJ) of India | Like most basal sauropods, this animal is very much like the larger prosauropods. |
Sauropoda i.s.: Okay, here's the big stuff.
A ?Kimmeridgian (?LJ) cervical from Yemen appears to belong
to a sauropod, and other Kimmeridgian remains in Argentina are from an
undescribed basal sauropod.
| Taxon or Taxa: | Time/Place: | Comments: | |
| Aepisaurus elephantinus (N.D.) Gervais, 1852 | Albian (EK) of France | This dubious sauropod may be a brachiosaurid, as its type humerus is apparently not either from a camarasaurid or titanosaurian, two earlier popular choices. It is sometimes misspelled Aepysaurus. | |
| Algoasaurus bauri (N.D.) Broom, 1904 | mid Tithonian-early Valanginian (LJ-EK) of South Africa | Based on poor remains rescued from pulverization into bricks, this animal could be a camarasaurid, titanosaurid, or diplodocid. | |
| Archaeodontosaurus descouensi Buffetaut, 2005 | Bathonian (MJ) of Madagascar | This sauropod is based on a lower jaw that deepens toward the front, as in derived sauropods, yet retains "prosauropod" (whatever prosauropod is now)-like teeth, with features like serrations. The describer notes that it is more common for basal sauropods to do the opposite, i.e. have shallow, "prosauropod"-like lower jaws but spoon-shaped sauropod teeth. This animal raises some questions about the diversity of Madagascar sauropods at that time, the describer stating that it differs from Lapparentosaurus and the "Bothriospondylus" complex, and that at least two sauropods are known from the deposits. | |
| ?Arkharavia heterocoelica (N.D.) Alifanov and Bolotsky, 2010 | ?late Maastrichtian (LK) of Russia | Arkharavia, known from caudals and teeth, was described as a sauropod similar to Chubutisaurus, but the holotype can only be assigned to Sauropoda with some charity, and other referred verts are hadrosaurid instead (probably from Olorotitan or Kundurosaurus, from the same site). | |
| Asiatosaurus mongoliensis (N.D.) Osborn, 1924 (?Euhelopus) | ?Valanginian-Albian (EK) of Mongolia and ?China | This tooth taxon may be the same as Euhelopus or a close relative. An Euhelopus-like vertebra from China has been tentatively referred to the type, without justification in my opinion. | |
| "Asiatosaurus" kwangshiensis (N.D.) Hou, Yeh, and Zhao, 1975 | ?Barremian-?early Aptian (EK) of China | Just like Asiatosaurus proper, this is a dubious sauropod based on teeth. | |
| "Bashunosaurus kaijiangensis" (N.N.) Kuang, 1996 vide Li, Zhang, and Cai, 1999? | ?MJ-?LJ of China | This sauropod was described as a camarasaurid, which leaves open the possibility it is actually an "euhelopodid." It's unclear exactly what the nomenclatural history for this name is. | |
| Blikanasaurus cromptoni Galton and Heerden, 1985 | early Norian (LTr) of Lesotho | This large basal sauropod is based on a partial hindlimb (minus femur) including a partial foot showing four stout metatarsals. This supports the idea that sauropod feet shortened for strength, and the fifth metatarsal "met them in the middle," so to speak. It has been traditionally considered a prosauropod. | |
| Bothriospondylus suffossus (N.D.) Owen, 1875 | early Kimmeridgian (LJ) of England | Based on vertebrae and not particularly well known, this sauropod has historically been regarded as close to Brachiosaurus (well, since there have been brachiosaurids, of course). | |
| "Bothriospondylus" elongatus (N.D.) Owen, 1875 | Berriasian-Valanginian (EK) of England | This indeterminate sauropod is based on a vertebra. | |
| "Brachiosaurus" nougaredi (?N.D.) Lapparent, 1960 | LJ of Algeria | The sparse remains of "B." nougaredi, a sacrum and forelimb and hindlimb material, are not enough to tell us if it really belongs in Brachiosaurus, or even if the remains all belong to the same kind of animal, especially when most of the remains were never collected or are lost. However, while perhaps not diagnostic in the classic sense, it is certainly distinctive, with a whopping huge sacrum. | |
| Bruhathkayosaurus matleyi (N.D.) Yadagiri and Ayyasami, 1989 | Coniacian (LK) of India | Bruhathkayosaurus is named from an ilium over 1.2 meters long, with some limb bones and et cetera referred. It was first described as a giant theropod, but the preservation is so poor other workers are unsure if the limb bones may not be petrified wood. Given the size, the time, and the place, it's most likely a titanosaurian of some sort, but little has been published beyond the initial description and its very unhelpful line drawings. | |
| "Campylodon" ameghinoi (N.D.) Huene, 1929 (Campylodoniscus) | late Cenomanian-early Turonian (LK) of Argentina | Also known as Campylodoniscus (Kuhn, 1961), this sauropod, based on an unusual maxilla, may be a late-surviving basal titanosaurian. It has teeth intermediate between basal sauropod spatulate teeth and titanosaurid pegs. | |
| Cardiodon rugulosus (?N.D.) Owen, 1844 | late Bathonian (MJ) of England | Based on a tooth, this taxon has long been referred to Cetiosaurus, which may be true but cannot be proven because no definite skull material has been found for European Cetiosaurus. In fact, recent study suggests that Cardiodon is not referable to Cetiosaurus and may represent a valid taxon based on its unique form. It could be a turiasaurian. | |
| "Cetiosaurus": | "C." longus (N.D.) Owen, 1842 | Kimmeridgian-Tithonian (LJ) of England | Occasionally referred to Cetiosauriscus, this is a dubious sauropod based on dorsal and caudal centra. |
| "C." medius ("type") (N.D.) Owen, 1842 | Bajocian (MJ) of England | Although this species has been known to science for longer than any other sauropod species, it is still poorly known (that's what being based on lousy material will do to you). At one time it was though to be a marine reptile. As it is a dubious type species for a well-known genus, and because of other taxonomic problems (see Pelorosaurus), a petition will be made to make C. oxoniensis the type species (and so I'm going to pretend it's already been done, so don't tell anyone, okay?). | |
| "C." mogrebiensis Lapparent, 1955 | late Bathonian (MJ) of Morocco | "C." mogrebiensis is known from the remains of three individuals. A nearly-complete skeleton referred to it actually belongs to the basal brachiosaurid Atlasaurus. It has been little studied, and there's no real reason to assign it to Cetiosaurus. | |
| Chiayusaurus lacustris (N.D.) Bohlin, 1953 (?Asiatosaurus or Mamenchisaurus) | Oxfordian-Kimmeridgian (LJ) or ?Barremian-Albian (EK) of China | Chiayusaurus may well be congeneric with another dubious sauropod tooth taxon, Asiatosaurus, or even Mamenchisaurus. Older references may list this animal as Chiayüsaurus, but such special symbols are now prohibited from official taxonomic use. | |
| "Chiayusaurus" asianensis (?N.D.) Lee, Yang, and Park, 1997 | Aptian-Albian (EK) of South Korea | This is a poorly-known EK sauropod. Like the genus to which it was assigned, it is based on a spatulate tooth. | |
| Chuanjiesaurus anaensis Fang, Pang, Lu, Zhang, Pan, Wang, Li, and Cheng, 2000 | early MJ of China | This large, newly-described taxon was referred to as a cetiosaurid. It is known from postcranial remains, including a scapula, humerus, numerous caudals, and forked chevrons. Remains from possibly three individuals are known, the largest possibly 25 meters in length (quite long for an early sauropod). | |
| Clasmodosaurus spatula (N.D.) Ameghino, 1899 | Santonian-early Campanian (LK) of Argentina | This is an indeterminate sauropod based on teeth. | |
| "Dachongosaurus yunnanensis" (N.N.) Zhao, 1983 | Sinemurian (EJ) of China | Also known as "Dachungosaurus", this informally-named basal sauropod is known from at least a partial articulated skeleton. | |
| "Damalasaurus magnus" (N.N.) Zhao, 1983 (species through Zhao, 1986) | EJ of China | Remains for "Damalasaurus" include at least a rib. Species "D. laticostalis" is apparently the same as "D. magnus", and may be the preferred name (at least as of 2006). | |
| Datousaurus bashanensis Dong and Tang, 1984 | Bathonian-Callovian (MJ) of China | Datousaurus is much more rare than its contemporaries Shunosaurus and Omeisaurus. In many ways it converges on the condition one would expect for a basal diplodocoid, but a possible skull is very robust and un-diplodocoid. For a basal sauropod, it is rather long, on the order of 50 feet long. | |
| Eomamenchisaurus yuanmouensis Lü J., Li T., Zhong S., Ji Q., and Li S., 2008 | MJ of China | Described as a mamenchisaurid related to M. huchuanensis and youngi, and Chuanjiesaurus, this sauropod is known from vertebral, girdle, and limb remains. | |
| Gigantosaurus megalonyx (N.D.) Seeley, 1869 | Kimmeridgian (LJ) of England | This is an indeterminate sauropod, possibly the same as others from the same time and area. It is not to be confused with the carcharodontosaurid theropod Giganotosaurus. | |
| "Hisanohamasaurus" (N.N.) Lambert, 1990 | LK of Japan | This undescribed sauropod is known from narrow-crowned teeth. | |
| "Iguanodon" praecursor (N.D.) Sauvage, 1876 | Kimmeridgian (LJ) of France | Like Gigantosaurus, this indeterminate sauropod may be the same as others from the same time and place. | |
| Klamelisaurus gobiensis Zhao, 1993 | mid Bathonian-late Callovian (MJ) of China | This sauropod converges in some way with the brachiosaurids. It has also been suggested as the adult form of Bellusaurus. | |
| Kunmingosaurus wudingensis Zhao vide Dong, 1992 | Hettangian (EJ) of China | Extremely obscure (it may technically be described, but that hasn't made anyone notice it), Kunmingosaurus is based on a partial skeleton centered on the pelvis and legs, with a couple of dentaries referred for good measure. | |
| "Lancanjiangosaurus cahuensis" (N.N.) Zhao, 1983 | MJ of China | This sauropod is supposedly known from cranial and limb fossils. "Lanchanjiangosaurus" is another spelling. "Lancangosaurus", usually thought to be an early name for Datousaurus, appears to be an older variant of "Lancanjiangosaurus"; the latter is preferred here as having wider usage, but with these informal names, it doesn't really matter. | |
| Macrurosaurus semnus (N.D.) Seeley, 1876 | Barremian (EK) of England | Macrurosaurus is based on caudal remains. Some of the assigned remains of this animal, but apparently not the type, pertain to a titanosaurian. | |
| Mamenchisaurus constructus Yang, 1954 | Oxfordian (LJ) of China | This sauropod is famous for its extremely long neck. Ironically, the species that was best known for this, "M." hochuanensis, exhibits many differences from the type and probably doesn't belong. That said, this animal is still very long-necked. A number of N.N. species have been referred to this taxon, but as they would tend to increase the clutter here for no good reason, I have left them out. The type species is known to have had forked chevrons and bifid presacral neural spines. | |
| "Mamenchisaurus": | "M." jingyanensis Zhang, Li, and Zheng, 1998 | ?Oxfordian (?LJ) of China | This taxon appears to be different from both Mamenchisaurus and "M." hochuanensis; among other differences, its dorsal vertebrae are procoelous, while those of the other two taxa mentioned are opisthocoelous. The hollows in its vertebrae are poorly developed, suggesting that this is a rather basal sauropod. |
| "M." anyuensis He, Yang, Cai, Li, and Liu, 1996 | Oxfordian (LJ) of China | Another day, another possible species of Mamenchisaurus. | |
| "Megacervixosaurus tibetensis" (N.N.) Zhao, 1983 | LK of China | This is an undescribed sauropod known from, unsurprisingly, cervicals. It might be a titanosaur. | |
| "Microdontosaurus dayensis" (N.N.) Zhao, 1983 | MJ of China | "Microdontosaurus" presumably is known from slender teeth, because it was assigned to an outdated superfamily for sauropods with such teeth. The name is already in use for an ichthyosaur, so another name would have to be used if this is ever published. | |
| Morinosaurus typus (N.D.) Sauvage, 1874 | Kimmeridgian (LJ) of France | Based on a tooth, this sauropod may be a titanosaurian, if it was correctly described and illustrated. | |
| Neosodon [no species name] (N.D.) Moussaye, 1885 | Kimmeridgian (LJ) of France | Based on a spoon-shaped tooth and usually tossed off as an indeterminate brachiosaurid, additional remains from the area suggest that it may have actually been a large "camarasaurid" or turiasaurian. However, since the type is only a tooth, it is hard to make the identification stick. It is sometimes listed as N. praecursor, with the teeth from "Iguanodon" praecursor assigned to it, but spoon-shaped sauropod teeth are very common. | |
| "Nemegtosaurus" pachi (N.D.) Dong, 1977 | early Maastrichtian (LK) of China | This indeterminate tooth taxon may be a nemegtosaurine. | |
| Ohmdenosaurus liasicus Wild, 1978 | mid Toarcian (EJ) of Germany | This basal sauropod is based on a partial hindlimb, which was once thought to belong to a plesiosaur. | |
| Omeisaurus junghsiensis Yang, 1939 |
Bathonian-Callovian (MJ) of China | Omeisaurus proper is known from several partial skeletons and a skull. What it constitutes has not yet been fully addressed. | |
| "Omeisaurus": | "O." changshouensis Yang, 1958 | Oxfordian (LJ) of China | This animal appears to be closer to Mamenchisaurus or "Mamenchisaurus" than to Omeisaurus. It has been a continual problem. |
| "O." jiaoi Jiang S., Li F., Peng G.-Z., and Ye Y., 2011 | Bathonian-Callovian (MJ) of China | "O." jiaoi is known from most of an articulated to associated skeleton including much of the back and tail, the pelvic girdle and hind limbs, half of the shoulder girdle, and most of the arms. The dorsals are opisthocoelous and have unbifurcated neural spines, and the anterior caudals are amphicoelous. | |
| "O." luoquanensis Li, 1988 | Bathonian-Callovian (MJ) of China | Like other species referred to Omeisaurus, this one may or may not belong here. | |
| "O." maoianus Tang, Jin, Kang, and Zhang, 2001 | Bathonian-Callovian (MJ) of China | This animal is from the same horizon as the type, and is known from a complete skull and partial postcranium including three cervicals, four dorsals, twenty-two caudals, most of the pelvic girdle, and most of the limbs. The presacral neural spines show no bifurcation. The skull is short, without much of a snout, and does not particularly resemble either that for "O." tianfuensis or O. junghsiensis. It may be more derived than the former. | |
| "Oshanosaurus youngi" (N.N.) Zhao, 1985 | EJ of China | "Oshanosaurus" is another in the parade of undescribed sauropods. Its claim to fame is that it was mistakenly thought to be a heterodontosaurid, due to proximity to a mention of former heterodontosaurid (current croc relative) Dianchungosaurus. | |
| Protognathosaurus oxyodon Olshevsky, 1991 (originally Protognathus oxyodon Zhang, 1988) | Bathonian-Callovian (MJ) of China | The original entry I had for this dinosaur
ran like this: "Here is my favorite pet hypothesis. Usually tossed off as an indeterminate
sauropod, I suspect that this animal may actually be a very early
therizinosaurian. It is based on a partial lower jaw with an unusual
downward bend. I have four reasons to suspect this is not a
sauropod: first, the bend of the jaw is unlike the bend in any known
sauropod, but compares well with known therizinosaurs (sauropod jaws
sometimes have a downward angle to the bottom, but never the top); second,
there are an unusually high number of teeth for a sauropod, but not for a
therizinosaurian; third, the shape of the teeth compares well with both
prosauropod and therizinosaurian teeth; and finally, the teeth are largest
at the front of the jaw and then become much smaller, a therizinosaurian
characteristic. This animal could be
either a basal therizinosaurian or a bizarre, unusually primitive sauropod.
Note that I have been wrong before..." Well, I'm here to say that I'm very sure I was wrong after having seen a presentation on a new basal EK therizinosaurian (Falcarius) at SVP. Suffice it to say that basal therizinosaurians were similar to other maniraptorans, and were apparently not pure herbivores. I now support the second possibility, that of a bizarre early sauropod. The jaw is distinctive, but we need some more of the rest of the animal to do much with it. |
|
| Qinlingosaurus luonanensis Xue, Zhang, and Bi, 1996 | mid-late Maastrichtian (LK) of China | Little is yet known of this sauropod. It is based on some postcranial material (ilium, ischium, and 3 verts) that doesn't distinguish it well from other sauropods. | |
| "Rebbachisaurus" tamesnensis (N.D.) Lapparent, 1960 (?Jobaria) | Albian (EK) of Algeria, Niger, and Tunisia, or based on material from sometime between the Late Triassic and Aptian, perhaps MJ of Niger | An apparently common EK African sauropod, "R." tamesnensis may have been a camarasaurid, unlike true Rebbachisaurus. It is also rather dubious. In fact, it is usually now just chucked in with Jobaria, but we'll see. Or we won't. | |
| "Sugiyamasaurus" (N.N.) Lambert, 1990 | K of Japan | This is an informal name for a Japanese sauropod, based on teeth of the "broad-toothed" variety. | |
| Tienshanosaurus chitaiensis Yang, 1937 (?Euhelopus) | Oxfordian (LJ) of China | Known from a partial postcranial skeleton, Tienshanosaurus is the first Chinese sauropod known from good remains and still one of its more obscure. | |
| "Titanosaurus": | "T." montanus (N.D.) Marsh, 1877 (Atlantosaurus) | Kimmeridgian (LJ) of Colorado | This indeterminate sauropod is better known under Marsh's second name for it, Atlantosaurus, which makes a fair amount of cameos in old dinosaur books. It is based on remains that most likely belong to Apatosaurus or something very much like it. |
| "T." rahioliensis (N.D) Mathur and Srivastava, 1987 | Maastrichtian (LK) of India | This is an indeterminate sauropod based on teeth. | |
| Tonganosaurus hei Li K., Yang C., Liu J., and Wang Z., 2010 | EJ of China | Tonganosaurus was described as a mamenchisaurid, although I choose to remain agnostic on such matters until it is determined what precisely Mamenchisaurus is. Tonganosaurus is known from verts, pectoral and pelvic material, and bones from the fore and hindlimbs. It joins the small but growing roster of EJ sauropods. | |
| Ultrasaurus tabriensis (N.D.) Kim, 1983 | Aptian-early Albian (EK) of South Korea | Unlike what its name implies, this was actually a small sauropod. Its type partial humerus was originally identified as a lower arm bone, making it seem larger than it really is. Because it was named before "Ultrasaurus" macintoshi, that animal had to be renamed Ultrasauros. It didn't really matter in the end, though, because Ultrasauros turned out to be Supersaurus. | |
| Xianshanosaurus shijiagouensis Lü J., Xu L., Jiang X., Jia S., Li M., Yuan C., Zhang X., and Ji Q., 2009 | Cenomanian (LK) of China | Xianshanosaurus is as-yet obscure. From the abstract of the description, I suspect it was a relatively large sauropod. | |
| "Yibinosaurus zhoui" (N.N.) Ouyang vide Anonymous, 2001 | EJ of China | This will likely turn out to be a new basal sauropod. | |
| "Yizhousaurus sunae" (N.N.) Chatterjee, Wang T., Pan S. G., Dong Z., Wu X. C., and Upchurch, 2010 | Hettangian-Sinemurian (EJ) of China | "Yizhousaurus" is a basal sauropod known from a nearly complete skeleton and skull. Unusually, the name came out as a Geological Society of America presentation abstract; as an abstract, that doesn't really count (the accepted citation for Parksosaurus was a GSA abstract, but I guess it was okay because it was renaming an already-named species). | |
| Zigongosaurus fuxiensis Hou, Zhao, and Chu, 1976 (?Mamenchisaurus, "Mamenchisaurus", Omeisaurus, or "Omeisaurus") | Bathonian-Callovian (MJ) of China | As can be seen, this taxon has very little "job security." Because it is uncertain what the sauropods listed as possible synonyms constitute, this animal is retained. It is known from sediments that are roughly between the age of those Omeisaurus is found in and those Mamenchisaurus is found in. Its neural spines show some bifurcation, but apparently its chevrons were simple. At one time, it was suggested to be a brachiosaurid. | |
| Zizhongosaurus chuanchengensis Dong, Zhou, and Zhang, 1983 | Toarcian (EJ) of China | Not to be confused with Zigongosaurus, this is another poorly-known basal sauropod. | |
Gravisauria: Eusauropoda and a few of its closest friends, which have also been called "vulcanodontids" or "barapsaurids".
| Taxon or Taxa: | Time/Place: | Comments: |
| Vulcanodon karibaensis Raath, 1972 | Hettangian (EJ) of Zimbabwe | Named for shed teeth found along with the type that turned out to belong to a theropod, Vulcanodon is one of the earliest known sauropods and also one of the least derived. |
| Spinophorosaurus nigerensis Remes, Ortega, Fierro, Joger, U., Kosma, Marín Ferrer, Chiappe, Dantas, Escaso, Gasulla, López, Pomares, Ribeiro, Sanz, Tent-Manclús, Faust, Joger, H., Joger, J., Krüger, Mudroch, Rabe, Ritter, Sommer, Ide, and Maga, 2009 [the authorlist is a bit unusual; see the paper] | sometime between the Late Triassic and Aptian, perhaps MJ of Niger | Spinophorosaurus is one of those lucky
cases where you get an almost-complete articulated skeleton right off the
bat; what's missing is part of the skull, the sternal elements, the arms
and hands, and the feet. Another specimen fortuitously enough has a
good chunk of what's not represented in the type specimen's skull. The type is subadult, while the paratype is more fully grown. Enough of the bragging. Spinophorosaurus plots as the sister group to Eusauropoda. It's fairly late for such a basal taxon (although note that the age of the formation has not been pinned down, so it could be older or younger). Its most striking feature (pun not intended) is the presence of spikes, which are interpreted as coming from the tail. The arrangement may have been a stegosaurian-like set of four paired spikes. Unlike Shunosaurus, which also had spikes, there was apparently no bony tail club. |
| Tazoudasaurus naimi Allain, Aquesbi, Dejax, Meyer, Monbaron, Richir, Rochdy, Russell, and Taquet, 2004 | Pliensbachian-Toarcian (EJ) of Morocco | This basal sauropod is known from the remains of an adult and juvenile; the adult remains include partial skull and (rather oddly bent, unless the photograph is of the piece unrestored) lower jaw, and a variety of postcranial material. Among the basal sauropods, it is closest to Vulcanodon. It was a smallish sauropod, on the order of 9 m long. |
| Barapasaurus tagorei Jain, Kutty, Roy-Chowdhury, and Chatterjee, 1975 | Sinemurian-Pliensbachian (EJ) of India | Barapasaurus is known from the most material of any EJ sauropod, including the partial remains of several individuals (all missing head, hands, and feet). |
| Rhoetosaurus brownei Longman, 1925 | Bajocian (MJ) of Australia | Known from much of the rear end of a moderately-sized generalized sauropod (partial tail, partial pelvis, right leg, and other odds and ends), Rhoetosaurus is a rare Australian dinosaur, and one of its first-described examples. Researchers return to the type locality occasionally and find additional parts. |
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