Iguanodontia
Iguanodontia includes the "iguanodonts" and the hadrosaurids. There are two general types of iguanodonts; small and large. Small iguanodonts, including the dryosaurids, look not too dissimilar to hypsilophodonts. Large iguanodonts, like Iguanodon, typically show quadrupedal tendencies, and sometimes have well-developed spikes for thumbs. They have robust, subrectangular skulls, with some modest flaring of the beak that becomes the "duck-bill" hadrosaurids are so known for. Many iguanodonts have lost the first toe as well. The jaws also begin to show the dental batteries and spreading ability, which in function converge with the chewing ability of mammals, that hadrosaurids have fully developed. Some have prominent extensions of the neural spines, giving them buffalo-like hump-back looks, including an unnamed taxon from Utah, and Ouranosaurus.
<--Iguanodontia
`--+--Gasparinisaura
|--Tenontosaurus
`--+--Rhabdodontidae
| |--Mochlodon
| |--Rhabdodon
| `--Zalmoxes
`--+--?Talenkauen
`--+--?Anabisetia
|--Dryosauridae
| |--Dryosaurus
| |--Dysalatosaurus
| `--Valdosaurus
`--Ankylopollexia (refers to the big ol' spiked thumbs)
|--Camptosauridae
| `--Camptosaurus
`--Styracosterna
|--Theiophytalia
`--+--Dakotadon
`--+--Fukuisaurus
`--Iguanodontoidea (also Hadrosauriformes)
|--Dollodon
|--Iguanodon
|--Mantellisaurus
`--Hadrosauroidea
|--Ouranosaurus
`--+--Altirhinus
|--Jinzhousaurus
`--+--Equijubus
`--+--Penelopognathus
|--Probactrosaurus
|--"Probactrosaurus" mazongshanensis
`--+--Eolambia
`--+--Protohadros
`--+--Jeyawati
`--+--Shuangmiaosaurus
`--+--Tanius
`--+--Bactrosaurus
`--+--Gilmoreosaurus
`-->Hadrosauridae
Iguanodontia: Basal Iguanodontia is a home for misfits, it seems; there is the hypsilophodont-like Rhabdodon
and its allies, the "iguanodontid"-like Muttaburrasaurus,
and the long-tailed oddball Tenontosaurus, among others. The
ordering is a bit controversial, and goes back and forth with which taxon is put
at the base of Iguanodontia.
| Taxon or Taxa: | Time/Place: | Comments: | |
| Gasparinisaura cincosaltensis Coria and Salgado, 1996 | Santonian-early Campanian (LK) of Argentina | Gasparinisaura was described as a dryosaur-like small iguanodontian, although some recent analyses indicates that it was closer to or less derived than Hypsilophodon. It is known from over 15 individuals, including the type specimen, a mostly-complete juvenile skeleton. | |
| Tenontosaurus: Ostrom, 1970 | T. tillettorum (type) Ostrom, 1970 | Aptian-Albian (EK) of Montana, Oklahoma, Texas, Utah, and Wyoming | Long-tailed, tall-spined Tenontosaurus is known from dozens of specimens of all ages. It seems to have been a favorite food source for Deinonychus. At times classified as a hypsilophodontid, at times as an iguanodontid, it seems to be between the two. It probably was mostly quadrupedal. Other species may be found in the copious material referred to the type species. |
| ?T. dossi Winkler, Murray, and Jacobs, 1997 | late Albian (EK) of Texas | T. dossi is more basal than the type, and is pretty long, on the order of 7 to 8 meters (not that the type species is any slouch). It may end up in its own genus. | |
| Talenkauen santacrucensis Novas, Cambiaso, and Ambrosio, 2004 | Maastrichtian (LK) of Argentina | From a new locality in southwestern Patagonia, Talenkauen is an iguanodontian known from a skeleton minus the tail, part of the ischia and pubes, hands, and ulnae. Probably on the order of 4 m long, it looked a lot like a dryosaurid, with an elongate neck and smallish head, but with long slender arms and a robust prepubic process. There were also thin mineralized plates along the sides of the ribcage; these probably were typically cartilaginous, and their function is unknown. Similar plates are known for several hypsilophodont-grade ornithischians. | |
| Anabisetia saldiviai Coria and Calvo, 2002 | Cenomanian-early Turonian (LK) of Argentina | Anabisetia is a new iguanodontian, based on a partial skull, parts of four cervicals, four dorsals, four sacrals, a dozen caudals, most of both shoulders and arms, and most of a leg. At least three more individuals are known, each about two meters in length. It is more derived than Gasparinisaura. | |
Iguanodontia i.s.: Several unnamed iguanodontians are known from the Cedar Mountain Formation, including a possible hadrosaurian.
| Taxon or Taxa: | Time/Place: | Comments: | |
| Bihariosaurus bauxiticus (N.D.) Marinescu, 1989 | ?Berriasian-?Hauterivian (?EK) of Romania | This obscure animal is an iguanodontian, possibly close to Camptosaurus but based on scanty remains | |
| "Cetiosaurus" brachyurus (N.D.) Owen, 1842 | mid Valanginian (EK) of England | This is a dubious species of Cetiosaurus named from an iguanodont dorsal and caudal (well, there wasn't much to go on yet). Which iguanodont it was will probably remain a permanent question, since you've got at least Barilium, Hypselospinus, and the original Iguanodon to choose from. | |
| Cumnoria prestwichii Seeley, 1888 (originally Iguanodon prestwichii Hulke, 1880) | Kimmeridgian (LJ) of England | Cumnoria, more often known as Camptosaurus prestwichii, is quite obscure and somewhat more generalized than the more famous Morrison Formation genus, although it's got a partial skeleton and skull to its credit. | |
| ?"Gadolosaurus" (N.N.) Saito, 1979 | ?K of Mongolia | This is an undescribed ?hadrosaurid, based on a (heavily-restored) juvenile specimen. It may be a juvenile of Arstanosaurus, or it may be closer to Camptosaurus or Iguanodon. | |
| ?"Gilmoreosaurus" atavus (N.D.) Nesov, 1995 | late Albian (EK) of Central Asia | This is either a very early hadrosaurid, or yet more basal iguanodontian, based on teeth. | |
| "Iguanodon": | "I." anglicus (N.D.) Holl, 1829 | mid Valanginian (EK) of England | As many of you probably know by now, the original type material for Iguanodon is pretty dodgy as these things go (some teeth), so the genus was conserved and I. bernissartensis made the type species. This leaves the teeth and postcranial bones referred to "I." anglicus in the lurch, as they probably aren't Bernissart Iguanodon, and nobody has really looked at them much in the last 150 years. They may come from either or both of Barilium and Hypselospinus (see below). |
| "I." orientalis (N.D.) Rozhdestvensky, 1952 | Aptian-Albian (EK) of Mongolia | "I." orientalis is based on a maxilla, ribs, and a scapula. For a while, the skull now named Altirhinus was thought to be a specimen, but obviously no longer is. It may be a specimen of I. bernissartensis (if its name was Marco Polo, I suppose), but the scrappy remains are not notably different from those of most dinosaurs of the iguanodontian persuasion. | |
| "I." ottigeri (N.D.) Galton and Jensen, 1979 | early Aptian (EK) of Utah | Based on teeth, this dubious Iguanodon species is sometimes seen named in connection with an undescribed high-spined iguanodontian from the same time and place. | |
| Macrogryphosaurus gondwanicus Calvo, Porfiri, and Novas, 2007 | Turonian-early Coniacian (LK) of Argentina | Macrogryphosaurus is a basal iguanodontian akin to Talenkauen (even including those odd lateral plates on the rib cage), known from postcranial remains. | |
| Planicoxa: DiCroce and Carpenter, 2001 |
P. venenica (type) DiCroce and Carpenter, 2001 | mid Aptian (EK) of Utah | A new Cedar Mountain Formation iguanodontian, Planicoxa (with its interesting name, to me reminiscent of a plant) is known from an ilium, vertebrae, and partial fore and hind limbs, belonging to at least two individuals. |
| P. depressa (Gilmore, 1909 [originally Camptosaurus] | Barremian (EK) of South Dakota | Long known as Camptosaurus depressus, the species of Camptosaurus that no one ever talked about (it seems that no one ever talks about C. prestwichii [Cumnoria], either, but it at least has the "English camptosaur" point in its favor), it is more similar to Planicoxa in the low depressed form of its ilium than it is to Camptosaurus. It is known only from the same site as the type specimen of Hoplitosaurus. | |
Rhabdodontidae: Rhabdodontids were not unlike robust, oversized hypsilophodonts, with deep skulls and jaws that were distinctly triangular in top view.
| Taxon or Taxa: | Time/Place: | Comments: | |
| Mochlodon suessi (Bunzel, 1871 [originally Iguanodon]) | ?early Campanian (LK) of Austria | Recent research suggests that Mochlodon is a separate taxon from Rhabdodon. Otherwise, it's not too dissimilar from the other rhabdodontids. | |
| Rhabdodon priscus Matheron, 1869 | Maastrichtian (LK) of France and Spain | Rhabdodon has gone through a complicated taxonomic
history. It may be found in older works under the name Mochlodon
Seeley
1881. Rhabdodon material is fairly common, but disarticulated. In the past, it has been regarded as an iguanodontid or a hypsilophodontid, although it now seems to be closest to animals like Tenontosaurus. A species named R. septimanicus Buffetaut and Le Loeuff 1991 is here tentatively regarded as a synonym of the type species. |
|
| Zalmoxes: Weishampel, Jianu, Csiki, and Norman, 2003 |
Z. robustus (type) (Nopcsa, 1899 [originally Mochlodon]) | early-mid Maastrichtian (LK) of Romania | Zalmoxes provides a new genus for the suddenly prolific rhabdomorphs, as they are sometimes known informally. No, the name of the second species is not a typo, but if you were ever playing Paleo Scrabble, it would be a good name to remember to get rid of the "q" (heck, if you were playing some renegade version where you got a couple dozen letters, adding the genus name would probably win the game outright. Or, you could just skip the species name and use Zalmoxes.). |
| Z. shqiperorum Weishampel et al., 2003 | early-mid Maastrichtian (LK) of Romania | ||
Dryosauridae: Dryosauridae is composed of several closely-knit taxa, which broadly resemble large hypsilophodonts. Annoyingly, there is a group of crocodilians known as dyrosaurids; be careful not to confuse them!
| Taxon or Taxa: | Time/Place: | Comments: |
| Dryosaurus altus Marsh, 1894 (originally Laosaurus altus Marsh, 1878) | Kimmeridgian (LJ) of Colorado, Utah, and Wyoming | Dryosaurus is known from remains including one virtually complete skeleton, and juvenile material. Like Dysalatosaurus, its skull does not completely encircle the two nostril openings. Interestingly, adults are not yet known for either taxon. |
| Dysalotosaurus lettowvorbecki Virchow, 1919 | late Kimmeridgian (LJ) of Tanzania | Its specific name coming from a WWI German general's name, and its generic name meaning "uncatchable lizard," it seems a political message is concealed within the name. Dysalatosaurus is known from a large amount of disarticulated material. It has long been referred to approximate contemporary Dryosaurus. |
| Valdosaurus canaliculatus Galton 1977 (originally Dryosaurus canaliculatus Galton, 1975) | Barremian (EK) of England | This taxon is known mostly from limb and jaw material. What is known links it with Dryosaurus. A fair amount of referred material is known, mostly for the hindlimbs, hips, and skull. Some material suggests individuals on Iguanodon atherfieldensis-scale (indeed, among material simply referred to Iguanodon may be iganodontoids, camptosaurs, and "rhabdomorphs") . |
Dryosauridae i.s.:
| Taxon or Taxa: | Time/Place: | Comments: |
|
Callovosaurus leedsi Galton, 1980 (originally Camptosaurus leedsi Lydekker, 1889) |
mid Callovian (MJ) of England | Once thought to be a hypsilophodont or camptosaurid, this basal iguanodontian is known from a partial femur. Its age makes it important in dating the earliest known extent of Iguanodontia. |
| Elrhazosaurus nigeriensis Galton, 2009 (originally Valdosaurus nigeriensis Galton and Taquet, 1982) | late Aptian (EK) of Niger | Elrhazosaurus is known from femora of the dryosaurid type. |
| Kangnasaurus coetzeei (N.D.) Haughton, 1915 | EK of South Africa | This is an indeterminate dryosaurid, based on teeth. Dryosaurid postcranial material is known from the area, and may belong to it. |
Camptosauridae: Camptosauridae, like Iguanodontidae, may not really exist as a monophyletic group.
| Taxon or Taxa: | Time/Place: | Comments: | |
| Camptosaurus: Marsh, 1885 | C. dispar (Marsh, 1879 [originally Camptonotus]) | Kimmeridgian (LJ) of Colorado, Oklahoma, Utah, and Wyoming | C. dispar is known from dozens of specimens. Although often thought of as an archetypical Morrison dinosaur, it is rather rare. Also, the trademark rectangular skull actually was a composite of actual Camptosaurus material, Theiophytalia kerri's skull (thought to be from the Morrison, but actually Early Cretaceous in age), and some jiggering to get the sections to fit. New skull material gives it a more triangular, basal look, something a bit more like Dryosaurus, and with long thin palpebrals. |
| C. aphanoecetes Carpenter and Y. Wilson, 2008 | Tithonian (LJ) of Utah | With the 2006 publication of a Camptosaurus skull that contained less than 50% accidentally reprocessed Theiophytalia, and the concurrent renovation of the Carnegie's dinosaur exhibits, the Dinosaur National Monument specimens of Camptosaurus were reevaluated by Carpenter and Wilson and found to differ from C. dispar in a variety of proportions. | |
Camptosauridae i.s.:
| Taxon or Taxa: | Time/Place: | Comments: |
| ?Brachyrophus altarkansanus (N.D.) Cope, 1878 (?Camptosaurus) | Kimmeridgian (LJ) of Colorado | Originally considered a sauropod, Brachyrophus may well be the same animal as Camptosaurus. Interestingly, if it is, its name predates that of Camptosaurus, making it technically the senior synonym, although the fact that it has been generally ignored while Camptosaurus has become a well-known name means that it is likely any attempt to change the name will be rejected. Incidentally, the only reason I have it here, instead of Ornithischia i.s. or something like that, is because of the likelihood it is the same as Camptosaurus. |
| Draconyx loureiroi Mateus and Antunes, 2001 | late Kimmeridgian-early Tithonian (LJ) of Portugal | Recently named from the partial remains of a single individual, mostly pertaining to the limbs (best material from the hindlimbs), Draconyx is a new member of the Lourinha fauna, related to Camptosaurus. |
Styracosterna:
| Taxon or Taxa: | Time/Place: | Comments: |
| Theiophytalia kerri Brill and Carpenter, 2006 | Aptian-Albian (EK) of Colorado | The so-called "Garden of the Gods" iguanodont, Theiophytalia is based on a partial skull (muzzle, cheek, and caudal half of the lower jaw, essentially) that seems akin to Dakotadon and somewhere between Camptosaurus (which it used to stand in for, until it was realized that the skull wasn't from the Morrison but from younger rocks) and Iguanodon. |
| Dakotadon lakotaensis Paul, 2008 (originally Iguanodon lakotaensis Weishampel and Bjork, 1989) | mid Barremian (EK) of South Dakota | Dakotadon is a much better supported North American possible Iguanodon species than "I." ottigeri, albeit without the cool "high-spined iguanodont" lurking in the background. It is based on a partial skull that, like Theiophytalia, seems to speak of an intermediate point betwixt Camptosaurus and Iguanodon. |
| Fukuisaurus tetoriensis Kobayashi and Azuma, 2003 | late Hauterivian-Barremian (EK) of Japan | Having been first referred to informally back in 1990, Fukuisaurus finally joins the ranks of the described. It is based on cranial material, including a deep lower jaw, and seems to have secondarily lost the cranial flexibility of other derived ornithopods, although it's hard to imagine why, given the usefulness of its proposed function providing an analogue to chewing. |
Styracosterna i.s.:
| Taxon or Taxa: | Time/Place: | Comments: |
| Barilium dawsoni Norman, 2010 (originally Iguanodon dawsoni Lydekker, 1888) | mid Valanginian (EK) of England | Barilium was a large (8 m scale in length) and relatively basal iguanodontian. It is known from several specimens and a fair amount of the skeleton, excepting the arms. |
| Hypselospinus fittoni Norman, 2010 (originally Iguanodon fittoni Lydekker, 1889) | mid Valanginian (EK) of England | Hypselospinus, a contemporary of Barilium, was smaller and less robust, with tall slender neural spines and heavily built arms. Included is Iguanodon hollingtoniensis, known from much of a skeleton. |
| Muttaburrasaurus langdoni Bartholomai and Molnar, 1981 | late Albian (EK) of Australia | Muttaburrasaurus is one of Australia's better-known dinosaurs. Superficially, it is very much like advanced iguanodontids, but in many ways it is actually very basal, and possibly is related to the hypsilophodont Atlascopcosaurus. It has a prominent, unusual rise on the nose, and has very odd teeth. Some researchers have suggested it was partially carnivorous. |
| Owenodon hoggii Galton, 2009 (originally Iguanodon hoggii Owen, 1874) | mid Berriasian (EK) of England | For a long time considered to be the earliest occurrence of Iguanodon, and then more recently thought to pertain to Camptosaurus, Owenodon appears to be somewhere between Camptosaurus and Lurdusaurus. It is based on a dentary. |
Iguanodon and Hadrosauroidea: So, what's the difference between
Iguanodontoidea
and Hadrosauroidea? The definition for Hadrosauroidea is written to
specifically exclude Iguanodon bernissartensis, while that of
Iguanodontoidea
is written to include it. Why do I have both in the diagram? When I
started, I was using Iguanodontoidea, but by the end of the first decade of the
2000s Hadrosauroidea had entered wider usage. Hadrosauriformes is the same
thing as Iguanodontoidea, just without a potentially-misleading suffix (all of
these "-oideas").
Many of these basal hadrosauroids were once classified as
iguanodontids,
but while they are similar to Iguanodon, they are not in the same family; rather,
they are part of the line that ends up as hadrosaurids. Many derived
hadrosauroids outside of the hadrosaurids (i.e. Bactrosaurus) were once
classified as hadrosaurids, but have since been pushed out of that club.
| Taxon or Taxa: | Time/Place: | Comments: | |
| Dollodon bambingi Paul, 2008 | Barremian-early Aptian (EK) of Belgium | This is the gracile Bernissart iguanodont, long known as I. mantelli or I. atherfieldensis. Gracile like Mantellisaurus, it is however not the same animal, as had been thought. | |
| Iguanodon bernissartensis Mantell, 1825 (type species after Boulenger, 1881) | Barremian-early Aptian (EK) of Belgium | This is going to be a little awkward, since
there's a decent chance that most or all of the Iguanodon species
deserve to be together as Iguanodon, and a decent chance that they
don't, but we're all used to talking about them together. For
safety, after removing I. atherfieldensis into Mantellisaurus,
I decided to pull the other species out into Iguanodontia i.s..
Honestly, when was the last time you cared about I. fittoni or I.
dawsoni? The way I'll think about it is that I.
atherfieldensis was due to be split by someone, Mantellisaurus
is just as good a name as any to recognize that, and Iguanodon should
just be thought of as the famous I. bernissartensis material. Iguanodon was one of the three founding members of Sir Richard Owen's Dinosauria. Originally, it was restored somewhat like the iguana it was named for, as a giant lizard with a horn on its nose (which turned out to be a thumb spike). Then, after the Belgian finds of the late 1800s, a tripodal kangaroo-like posture came to be favored. Finally, it has become a quadruped since about the 1980s, most notably in Disney's Dinosaur. Although Iguanodon gets the fame for being a founding member, the original type material was pretty poor (teeth, if your memory serves you well), and given the potential for taxonomic weirdness, the name was recently shifted to a specimen of I. bernissartensis. I. bernissartensis is known from 26 skeletons and much partial and isolated material, including the famous Bernissart quarry, where at least 32 individuals were found. It is much more robust than Mantellisaurus and Dollodon, with long massive forelimbs (and a hadrosaurid-length body of 10 to up to 13 meters), while Mantellisaurus was much shorter, at around 6 to 7 meters in length, and likely a lower browser. Iguanodon is more common in Belgium, with Dollodon the local gracile genus, but the gracile Mantellisaurus is more common in England. It has occasionally been suggested that Mantellisaurus and I. bernissartensis are the two sexes of one species, but this now appears to be an untenable position. |
|
| Mantellisaurus atherfieldensis Paul, 2006 (originally Iguanodon atherfieldensis Hooley, 1924) | early Aptian (EK) of England | To continue from above: this is the
"gracile (skinny, slim, fine-boned, what-have-you) Iguanodon".
A taxon named Vectisaurus
was once thought to be different because of its tall neural spines, but now appears
to be a specimen of this similarly tall-spined animal. Although I. atherfieldensis probably needs its own generic name, precisely which name to use is a frustrating matter: literally, it is the youngest of several probable synonyms, which is not supposed to happen. Apparently, several goes were made at naming the "gracile iguanodont", including I. mantelli (Meyer, 1832; a misinterpretation, as it was based on the same material as I. anglicus, but became used for the gracile Bernissart iguanodont now known as Dollodon), Heterosaurus neocomiensis (Cornuel, 1850), Sphenospondylus gracilis (Lydekker, 1888), and the above-mentioned Vectisaurus valdensis (Hulke, 1879), and I. atherfieldensis is just based on the best material. We'll see how that flies in the next few years (this must be like waiting for someone to challenge the constitutionality of a law), but Mantellisaurus is a good start. |
|
| Ouranosaurus nigeriensis Taquet, 1976 | late Aptian (EK) of Niger | Ouranosaurus is most famous for its well-developed ridge-back, much like that of some spinosaurids. It had a sort of knob right in front of the eyes. A couple of good skeletons are known. | |
| Jinzhousaurus yangi Wang and Xu, 2001 | early Aptian (EK) of China | A Yixian dinosaur, although not from the same level that has produced so many feathered finds, Jinzhousaurus is a large (~7 m long) iguanodontian based on material including an almost complete skeleton and skull that show an odd mix of basal and derived characters. It apparently is not as close to hadrosaurids as Probactrosaurus. | |
| Altirhinus kurzanovi Norman, 1998 | late Aptian-early Albian (EK) of Mongolia | Based on material once referred to the now-defunct taxon Iguanodon orientalis, this hadrosauroid had a prominent nasal arch. | |
| Equijubus normani You, Luo, Shubin, Witmer, Tang Z., and Tang F., 2003 | Albian-?Cenomanian (EK-?LK) of China | Very little has yet been revealed on this possible basal "hadrosaurian," but I could not let such a great name go by unnoticed. A short-snouted skull and partial postcranial remains are known. It is said to be similar to Altirhinus and Jinzhousaurus. | |
| Penelopognathus weishampeli Godefroit, Li, and Shang, 2005 | Albian (EK) of Mongolia | This hadrosauroid, based on a long, low right dentary (most hadrosauroids tend to have short, robust lower jaws), may be more derived than Altirhinus but less than Probactrosaurus based on dental characters. | |
| Probactrosaurus gobiensis Rozhdestvensky, 1966 (including P. alashanicus Rozhdestvensky, 1966) | Barremian-Aptian (EK) of China | Probactrosaurus is the sort of animal that would be expected as the ancestor of the hadrosaurids (made explicit in the name: "before Bactrosaurus"), if the material has been interpreted correctly. It has a plain iguanodont skull, without crests or knobs. | |
| "Probactrosaurus" mazongshanensis Lu, 1997 | Barremian (EK) of China | Based on fragmentary remains, this species seems to be closer to Altirhinus than to Probactrosaurus. | |
| Eolambia caroljonesa Kirkland, 1998 | latest Albian-earliest Cenomanian (EK-LK) of Utah | Once thought as to be the earliest known hadrosaurid (but not the most generalized-Protohadros had that honor), this was originally going to be called "Eohadrosaurus caroljonesi" before its supposed lambeosaurine affinities were clear (more recent work suggests it is a more basal iguanodontian). Several individuals are known, and their remains make up most of a skeleton. | |
| Protohadros byrdi Head, 1998 | Cenomanian (LK) of Texas | This recently-described dinosaur, based on a skull and some postcranial bits, was first described as the most basal hadrosaurid yet discovered, but it now appears to be a derived non-hadrosaurid iguanodontian. It has an unusually large lower jaw. | |
| Jeyawati rugoculus McDonald, Wolfe, and Kirkland, 2010 | middle Turonian (LK) of New Mexico | Jeyawati is a hadrosauroid that falls out somewhere between the "near-hadrosaurs" Probactrosaurus, Eolambia, and Protohadros, and the traditional "base of Hadrosauridae" characters Bactrosaurus, Telmatosaurus, and friends. It is known from a partial skull and skeleton, with the most obvious distinguishing feature being its knobbly postorbitals. | |
| Shuangmiaosaurus gilmorei You, Ji, Li J., and Li Y., 2003 | Cenomanian-Turonian (LK) of China | Described as the sister taxon to Hadrosauridae, Shuangmiaosaurus is based on partial remains of the upper and lower jaws. | |
| Tanius sinensis Wiman, 1929 | early-mid Maastrichtian (LK) of China | Considered at times to be congeneric with Tsintaosaurus, this obscure and mysterious hadrosauroid is known from ?three disarticulated specimens. At times it has been classified as a basal hadrosaurid, although it may have lambeosaurine affinities. | |
| Bactrosaurus johnsoni Gilmore, 1933 | early Maastrichtian (LK) of China | Bactrosaurus is pretty close to the dividing line between hadrosaurids and their more basal cousins, and is often described as a basal hadrosaurid. It is known from partial skeletons belonging to robust individuals. According to the common interpretation, at the time of describing an iguanodont-like skull got mixed in with the remains and made an animal with a classic lambeosaurine postcranium into a hadrosaurine. Later, it was discovered that two hadrosaurids were represented in the remains, one being the lambeosaurine Bactrosaurus, the other (and the owner of the skull) a hadrosaurine-like animal now named Gilmoreosaurus. However, some researchers are suggesting, based on further bonebed material, that Bactrosaurus was not chimeric and the flat skull actually a part of one individual (but still separated from Gilmoreosaurus), making this animal not a lambeosaurine, but a more basal hadrosauroid. | |
| Gilmoreosaurus: Brett-Surman, 1979 | G. mongoliensis (type) (Gilmore, 1933 [originally Mandschurosaurus mongoliensis]) | early Maastrichtian (LK) of China | Gilmoreosaurus was long confused with
the dubious hadrosaurid Mandschurosaurus. Some material once referred to
the early hollow-crested hadrosaurid Bactrosaurus, appears to actually belong to the contemporaneous Gilmoreosaurus
instead, which has a more gracile body. G. arkhangelski may be chimeric. |
| ?G. arkhangelskyi (?N.D.) Nesov, 1995 | late Turonian-Coniacian (LK) of Uzbekistan | ||
Hadrosauroidea i.s.:
| Taxon or Taxa: | Time/Place: | Comments: |
| Cedrorestes crichtoni Gilpin, DiCroce, and Carpenter, 2006 | early Aptian (EK) of Utah | Based on a sacrum, one complete ilium and a chunk of the other, a right tibia and 3rd metatarsal, ribs, and the inescapable ossified tendons always to be found with ornithopod skeletons consisting of more than bits and pieces, Cedrorestes is a contender for oldest basal hadrosaurid. This assessment is based on features of the ilium, especially the presence of an antitrochanter (a lateral knob on the caudal portion of the ilium that hadrosaurids have but their more basal cousins lack). I'm a little wary of basing classifications on one character, and of basalmost hadrosaurids (Eolambia and Protohadros, anybody?), so I'll put it here for the time being. |
| Glishades ericksoni Prieto-Márquez, 2010 | Campanian (LK) of Montana | Glishades is based on two partial premaxillae from the Two Medicine Formation, from a derived hadrosauroid, but not a true hadrosaurid. It may have been closely related to Bactrosaurus. |
| Jintasaurus meniscus You H. and Li D., 2009 | EK of China | Jintasaurus is known from a partial posterior skull from a derived non-hadrosaurid iguanodont, but there seems to be something odd going on with the definitions used in the description. It was described as the sister group to Hadrosauroidea, but that doesn't make sense: Hadrosauroidea is defined as all iguanodonts closer to Parasaurolophus than Iguanodon, which would kind of make Iguanodon the default sister group of Hadrosauroidea. It is also supposed to be more derived than other EK hadrosauriforms, but again by definition, the only hadrosauriform that is not a hadrosauroid is Iguanodon. This leaves me somewhat puzzled, but if you consider it as more derived than Protohadros, as the authors did, that at least puts you into familiar real estate. |
| Lanzhousaurus magnidens You, H., Q. Ji, and D. Li, 2005 | EK of China | As others have noticed, a Google search of Lanzhousaurus brings up a picture of a skeletal mount of a big ornithopod, certainly iguanodont-grade, as revealed by its astonishingly huge teeth (hadrosaurids have much smaller teeth). The mount is misleading, as only a lower jaw, teeth, some cervicals and dorsals, ribs, sternal elements, and both pubic bones are known. From what is known, a hadrosaurid length (~10 m, 33 ft) is suggested; a rather tubby mass of 5.5 metric tons is also suggested, which seems off given that most hadrosaurids of the same length are thought to have massed more like 2-3 metric tons. |
| Levnesovia transoxiana Sues and Averianov, 2009 | middle-late Turonian (LK) of Uzbekistan | One man's hadrosauroid is another man's basal hadrosaurid, depending on how the definitions are worked. Levnesovia was described as a basal hadrosauroid related to Bactrosaurus which, for the purposes of this site, puts it in Hadrosauridae. The two, possibly with Gilmoreosaurus, may represent an early radiation of hadrosaurids/oids. Material including cranial and vertebral fossils is known to date. |
| Lurdusaurus arenatus Taquet and Russell, 1999 | late Aptian (EK) of Niger | Having long gone under the informal name "Gravisaurus
tenerensis" (Chabli, 1988), this animal was one of the most heavily-built
iguanodontians. It has been compared to the prehistoric giant ground
sloth. Remains of two individuals of this unusual iguanodontian are known. It may have gotten to be around 9 meters long, with short hindlimbs and stout forelimbs armed with enlarged thumb spikes, making their arms like maces. |
| Nanyangosaurus zhugeii Xu, Zhao, Lu, Huang, Li, and Dong, 2000 | ?Albian (EK) of China | This animal, based on a partial postcranial skeleton, appears to be just basal to the true hadrosaurids. |
Hadrosauridae: Hadrosauridae is the group comprising the "duck-bills." It also is literally swamped in names given to indeterminate remains.
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