Abelisauridae
Abelisauridae is a clade of theropods that was first
described in the mid 1980s, from Abelisaurus and Carnotaurus, two
new LK species from Argentina. So far, they are known mostly from the late
Cretaceous of the Southern
Hemisphere. Many of them appear to have been fairly large. They were
the Gondwanan equivalent of the tyrannosaurids,
being the top predators (after the spinosaurids and
carcharodontosaurids were out of the way in the
early Late Cretaceous). However, they probably went about their business
in different ways than tyrannosaurids, given their less cursorial builds (abelisaurids
seem to have been longer and lower than tyrannosaurids, from what we currently
know), less robust skulls and teeth, and different prey items (Gondwanan
continents being overrun with titanosaurians and
with not much in the way of hadrosaurids, ankylosaurians,
or ceratopsians).
The traditional definition of Abelisauridae has Abelisaurus
off in a corner and a group of carnotaurines, which is a bit unfair since Abelisaurus
is only known from a partial skull. "Carnotaurines" are
traditionally known for very short arms practically lacking the forearm
segment, and for elaborate horns and knobs on the skull. Abelisaurid
skulls are known for having deep upper jaws with tall snouts, and slim lower
jaws. They also have "cheeks" expanded side to side, like those of
tyrannosaurids, perhaps conferring an additional degree of binocular vision
over predators with more narrow skulls.
Besides its two founding members, Abelisaurus and Carnotaurus,
Abelisauridae has become a home for a number of poorly-known theropods whose affinities
had previously been unknown. With the recent flood of good material from South
America and Madagascar, it is very likely that this group will continue to increase in size.
<--Abelisauridae
|--Kryptops
`--+--Rugops
`--+--Ekrixinatosaurus
`--+--+--Carnotaurus
| `--Ilokelesia
`--+--Indosaurus
|--Majungasaurus
`--Rajasaurus
Abelisauridae: This is a sort of theropod "rogue's gallery," where
the members are outlandish in appearance. Horns above the eyes (Carnotaurus
means "meat-eating bull"), extraordinarily deep skulls matched with slender
dentaries, tiny, seemingly useless arms and hands, and strange thickening of the skull
roof are things seen in some or all of these creatures. It is most probable that the
cranial ornamentation was used for display. "Carnotaurines" also have
L-shaped "wings" (sticking out and up on the long end of the L, with
the short end pointing forward) for transverse processes on the caudal
verts.
There are a number of poorly-known
abelisaurids described from India (Huene and Matley had a great big monograph
back in 1933) that have suffered greatly, first from initial misinterpretations
(and the fact that they weren't complete enough to draw good comparisons),
and later from specimen loss; a great deal of the convoluted taxonomy could
probably be cleaned up if the original specimens could be located. For
example, it is entirely within the realm of possibility that Indosaurus, Indosuchus,
and Compsosuchus are the same thing, and Lametasaurus
and Rajasaurus the same thing. It appears that there were at least
two large theropods, possibly three, in the Lameta, one represented by
Indosuchus, one represented by Carnotaurus-like Rajasaurus,
and a possible non-abelisaurid abelisauroid (suggested by some skull elements with
non-abelisaurid features).
An abelisaurid tooth, previously referred to Majungasaurus,
is known from the Maastrichtian (LK) of Egypt.
"Carnotaurines", particularly the distinctive namesake, have
become the go-to theropods when someone wants to depict a theropod that is not
either Tyrannosaurus, Velociraptor
(or another "raptor"), or Allosaurus.
| Taxon or Taxa: | Time/Place: | Comments: |
| Kryptops palaios Sereno and Brusatte, 2008 | Aptian-Albian (EK) of Niger | Kryptops is another rough-faced abelisaurid, with an upper jaw laced with vessel tracks and pits indicating some kind of covering that was firmly attached. It is one of the earliest described abelisaurids, and is known from a partial skeleton including a maxilla, verts and ribs, and a sacrum and partial pelvis, from a mature animal around the size of Majungasaurus. |
| Rugops primus Sereno, Wilson, and Conrad, 2004 | Cenomanian (LK) of Niger | Rugops is a hornless (no bony horns, at least) basal abelisaurid, known from a partial skull lacking the palate and most of the bones along the side of the face behind the eyes. This skull is distinguished by a row of depressions along the upper surfaces of the nasals, and by a small hole between the lacrimal, prefrontal, frontal, and postorbital (which is a little sketchy on absolute dimensions given that the postorbitals are AWOL). It has been suggested that these holes held blood vessels to supply a nonbony keratinous or fleshy ridge or crest system. I like the name; it is short and direct, like a good predator ought to be. |
| Ekrixinatosaurus novasi Calvo, Rubilar-Rogers, and Moreno, 2004 | late Cenomanian (LK) of Argentina | Ekrixinatosaurus (name means
"explosion-born lizard," referring to being discovered during a
blasting operation) is based on a fair amount of a skeleton belonging to a
single individual, about 6 m long from its skeletal restoration.
Both maxillae and dentaries, part of the braincase, a couple of cervicals,
many dorsals, the sacrum, anterior-middle caudals, most of the pelvis
(except for the distal ischia), ribs, most of one hindlimb, and the knee
of the other hindlimb were preserved. It has standard
"carnotaur" caudal process wings and pelvic form. It has a relatively bigger head than Carnotaurus (extrapolated skull-femur ratio 1.00 compared to 0.58), but is not particularly unique compared to other abelisaurids (some different proportions and holes and depressions on the verts, and so on). |
| Carnotaurus sastrei Bonaparte, 1985 | Maastrichtian (LK) of Argentina | This theropod is known from some of the best material of any dinosaur. Its type is a virtually complete skeleton and skull with extensive skin impressions, showing this animal to have had widely-spaced rows of small scutes, in addition to the bony horns which give this theropod its name. Its hands and arms were so small as to be virtually useless; for example, the lower arm bones (ulna and radius) were functionally part of the hand. Once dated to the "middle" Cretaceous, it is now known to be much younger. |
| Ilokelesia aguadagrandensis Coria and Salgado, 2000 | Turonian-Santonian (LK) of Argentina | This is a new, incomplete abelisaurid with a very interesting name, known from skull fragments, some vertebrae, and hindlimb material first found before at least 1991. It was initially proposed as a basal sister group to Abelisauridae+Noasauridae. The caudal verts' transverse processes are T-shaped. |
| Indosaurus matleyi Huene and Matley, 1933 (?Indosuchus) | Maastrichtian (LK) of India | I am sad to report that Indosaurus does not appear to have had horns after all, and could potentially be the same as Indosuchus (unfortunately the type material is lost). It was once identified as an allosaurid or "megalosaurid". |
| Majungasaurus crenatissimus Lavocat, 1955 (originally Megalosaurus crenatissimus Depéret, 1896; including Majungatholus atopus Sues and Taquet, 1979) | mid Maastrichtian (LK) of Madagascar | This theropod has been through a mess.
For a long time, there was Megalosaurus crenatissimus, or Dryptosaurus
crenatissimus, or Majungasaurus crenatissimus, a large
theropod. Then, there was Majungatholus, thought to be a large,
strange pachycephalosaurid. Then,
a recently-discovered
skull set the record straight partially by showing that Majungatholus
was really a theropod. Because the crenatissimus material
wasn't the greatest, it was left out of the party until there were enough
remains to show that indeed it was the same as Majungatholus and by
priority should be the name used. In one of those things that happen, "Megalosaurus" crenatissimus and Majungasaurus actually are based on different material. Depéret based his species on a collection of teeth, verts, and a phalanx, with no actual holotype, whereas Lavocat designated a Carnotaurus-like dentary from an immature individual as the neotype for Majungasaurus, while explicitly recognizing "M." crenatissimus and Majungasaurus to be the same thing. Among the many interesting skull features are the thickened nasals, a horn on the frontal, a parietal prominence, and pneumatic chambers. Remains from several individuals and size classes are known. Tooth markings on some remains suggest cannibalism. |
| Rajasaurus narmadensis Wilson, Sereno, Srivastava, Bhatt, Khosla, and Sahni, 2003 | Maastrichtian (LK) of India | Based on material including a skull with a low horn, hip material, caudals, and hindlimb material, this new large abelisaurid puts interesting questions to those who delight in taxonomy, given the tangled state of theropod taxonomy of the Lameta Formation of India. It may put to rest some of the controversy surrounding Lametasaurus, as its ilia are very similar to the ilia included in the material of the latter. |
Abelisauridae i.s.:
| Taxon or Taxa: | Time/Place: | Comments: |
| Abelisaurus comahuensis Bonaparte and Novas, 1985 | early Campanian (LK) of Argentina | Known only from a partial skull, Abelisaurus is usually regarded as closely related to Carnotaurus, although that may seem hard to believe based on the superficial aspects of the skulls (and some researchers have recently suggested that it may be a late-surviving carcharodontosaurid). It is one of my favorite big theropods; I like the no-nonsense look of the skull. |
| Aucasaurus garridoi Coria, Chiappe, and Dingus, 2002 | early Campanian (LK) of Argentina | This abelisaurid is based
on an almost complete skeleton back to the 13th caudal, including soft tissue impressions about
the hips. The animal is considered to be about 70% of Carnotaurus'
size, with a longer, lower skull with bumps instead of horns, but very
similar to the other animal. The
arms were also somewhat longer. It had been buried in a shallow lake; damage to the skull suggests it had been involved in a fight shortly before death. |
| Dryptosauroides grandis (N.D.) Huene and Matley, 1933 | Maastrichtian (LK) of India | This theropod is based upon six incomplete caudals that have no features that would allow it to be distinguished from the multitude of other dubious theropods from the Lameta Formation, or from Majungasaurus. It would have been large, though, on the order of Carnotaurus in size. |
| Indosuchus raptorius Huene, 1932 (?Compsosuchus, ?Indosaurus) | Maastrichtian (LK) of India | This theropod is based on currently absent skull material. It was a contemporary of
Rajasaurus and (possibly synonymous) Indosaurus,
but appears to be closest to Abelisaurus and Majungasaurus.
It was originally identified as a tyrannosaurid. Much new material has been assigned to it, making up most of a skeleton. This material includes a relatively long arm, especially when compared to Carnotaurus. Unfortunately, the fact that several other large theropods (possibly synonymous) are known from the same time and place means that this identification is not secure. |
| Lametasaurus indicus (N.D.) Matley, 1921 | Maastrichtian (LK) of India | This taxon was once considered to be an armored dinosaur, but now its remains are believed to belong to a theropod, a crocodilian, and an armored dinosaur, either a titanosaurian or ankylosaurian. In particular, the type is given as scutes (by some sources; other suggest differently), which are a mix of crocodilian and possible titanosaur and/or ankylosaurian armor; referred material includes ilia, a tibia, and a sacrum, which may belong to Rajasaurus. |
| Ornithomimoides mobilis (N.D.) Huene and Matley, 1933 | Maastrichtian (LK) of India | The specimens assigned to this species are inadequate for classification beyond Abelisauridae, and are essentially identical to those of Majungasaurus. It was based on five caudals (or six caudals). |
| ?Pycnomenosaurus nevesi Kellner and Campos, 2002 | ?Santonian (LK) of Brazil | This is a new abelisaurian from Brazil, apparently the first such named member of the clade from that country. It is based on rather scrappy remains, including a tibia, parts of a fibula and pubis, and a handful of verts, uncovered in the early 1950s, and may be an abelisaurid. |
| Quilmesaurus curriei Coria, 2001 | late Campanian (LK) of Argentina | Based on a small to medium sized femur and tibia, Quilmesaurus initially defied classification, but it now appears as though this animal probably was an abelisaurid (?"carnotaurine"). |
| Skorpiovenator bustingorryi Canale, Scanferla, Agnolin, and Novas, 2008 | late Cenomanian-early Turonian (LK) of Argentina | Skorpiovenator is named for the numerous scorpions inhabiting the dig site. It is based on a mostly complete skeleton lacking the shoulders and most of the arms (typical), the ends of the ischia and pubes, and about half of the tail. Unlike several other abelisaurids, the skull has no prominent bony ornamentation, but is rather deep and vaguely dog-like in profile (actually, now that I think about it, it's probably not all that far off from a complete Abelisaurus skull). The type individual was about 6 m long. |
| Vitakridrinda sulaimani Malkani, 2006 | Maastrichtian (LK) of Pakistan | Vitakridrinda was named for a partial skull and femoral remains, with some referred vertebrae. |
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